Adeoye, A., Jaramillo-Torres, A., Fox, S., Merrifield, D., Davies, S., 2016. Supplementation of formulated diets for tilapia (Oreochromis niloticus) with selected exogenous enzymes: Overall performance and effects on intestinal histology and microbiota. Animal Feed Science and Technology 100(215), 133-143. DOI: 10.1016/j.anifeedsci.2016.03.002
Arif, M., Alagawany, M., Abd El-Hack, M., Saeed, M., Arain, M., Elnesr, S., 2019. Humic acid as a feed additive in poultry diets: A review. Iranian Journal of Veterinary Research 20(3), 167-172.
Aristimunha, P.C., Mallheiros, R., Ferket, P.R., Cardinal, K.M., Moreira Filho, A.L.D.B., Santos, E., Cavalcante, D.T., Ribeiro, A.M.L., 2020. Effect of dietary organic acids and humic substance supplementation on performance, immune response and gut morphology of broiler chickens. Journal of Applied Poultry Research 29(1), 85-94. DOI: 10.3382/japr/pfz031
Costa, S.T.D., Sutili, F.J., Gressler, L.T., Loebens, L., Colpo, C.V., Lazzari, R., Baldisserotto, B., 2016. Growth of silver catfish (Rhamdia quelen) exposed to acidic pH at different humic acid levels. Ciência Rural 46(6), 1094-1099. DOI: 10.1590/0103-8478cr20151027
Forsatkar, M.N., Nematollahi, M.A., Amiri, B.M., Huang, W.-B., 2014. Fluoxetine inhibits aggressive behaviour during parental care in male fighting fish (Betta splendens, Regan). Ecotoxicology 23(9), 1794-1802. DOI: 10.1007/s10646-014-1345-0
Forsatkar, M.N., Nematollahi, M.A., Brown, C., 2017. Male Siamese fighting fish use gill flaring as the first display towards territorial intruders. Journal of Ethology 35(1), 51-59. DOI: 10.1007/s10164-016-0489-1
Forsatkar, M.N., Nematollahi, M.A., Rafiee, G., Farahmand, H., Lawrence, C., 2018. Effects of the prebiotic mannan‐oligosaccharide on feed deprived zebrafish: Growth and reproduction. Aquaculture Research 49(8), 2822-2832. DOI: 10.1111/are.13745
Gao, X., Zhang, H., Xu, Y., Ni, Q., Zhang, Y., Tan, H., Gu, C., 2022. Effects of humic acid on the nitrogen utilization efficiency and microbial communities in aquaponic systems. Aquaculture 547, 737475. DOI: 10.1016/j.aquaculture.2021.737475
Gao, Y., He, J., He, Z., Li, Z., Zhao, B., Mu, Y., Lee, J.-Y., Chu, Z., 2017. Effects of fulvic acid on growth performance and intestinal health of juvenile loach Paramisgurnus dabryanus (Sauvage). Fish & Shellfish Immunology 100(62), 47-56. DOI: 10.1016/j.fsi.2017.01.008
Kamali Najafabad, M., Imanpoor, M.R., Taghizadeh, V., Alishahi, A., 2016. Effect of dietary chitosan on growth performance, hematological parameters, intestinal histology and stress resistance of Caspian kutum (Rutilus frisii kutum Kamenskii, 1901) fingerlings. Fish Physiology and Biochemistry 42(1), 1063-1071. DOI: 10.1007/s10695-016-0197-3
Lichak, M. R., Barber, J. R., Kwon, Y. M., Francis, K. X., Bendesky, A., 2022. Care and use of Siamese fighting fish (Betta Splendens) for Research. Comparative Medicine 72(3), 169-180. DOI: 10.30802/AALAS-CM-22-000051
Lieke, T., Steinberg, C. E., Pan, B., Perminova, I. V., Meinelt, T., Knopf, K., Kloas, W., 2021. Phenol-rich fulvic acid as a water additive enhances growth, reduces stress, and stimulates the immune system of fish in aquaculture. Scientific Reports 11(1), 174-186. DOI: 10.1038/s41598-020-80449-0
Louvado, A., Cleary, D. F., Pereira, L. F., Coelho, F. J., Pousão-Ferreira, P., Ozório, R. O., Gomes, N. C., 2021. Humic substances modulate fish bacterial communities in a marine recirculating aquaculture system. Aquaculture 544, 737121. DOI: 10.1016/j.aquaculture.2021.737121
Lumactud, R. A., Gorim, L. Y., Thilakarathna, M. S., 2022. Impacts of humic-based products on the microbial community structure and functions toward sustainable agriculture. Frontiers in Sustainable Food Systems 6, 977121. DOI: 10.3389/fsufs.2022.977121
McDonald, S., Bishop, A.G., Prenzler, P.D., Robards, K., 2004. Analytical chemistry of freshwater humic substances. Analytica Chimica Acta 527(2), 105-124. DOI: 10.1016/j.aca.2004.10.011
Meinelt, T., Schreckenbach, K., Knopf, K., Wienke, A., Stüber, A., Steinberg, C. E., 2004. Humic substances affect physiological condition and sex ratio of swordtail (Xiphophorus helleri Heckel). Aquatic Sciences 66(2), 239-245. DOI: 10.1007/s00027-004-0706-9
Merrifield, D. L., Harper, G. M., Mustafa, S., Carnevali, O., Picchietti, S., Davies, S. J., 2011. Effect of dietary alginic acid on juvenile tilapia (Oreochromis niloticus) intestinal microbial balance, intestinal histology and growth performance. Cell and Tissue Research 344(1), 135-146. DOI: 10.1007/s00441-010-1125-y
Nur, F. M., Batubara, A. S., Fadli, N., Rizal, S., Siti-Azizah, M. N., Muchlisin, Z. A., 2022. Elucidating species diversity of genus Betta from Aceh waters Indonesia using morphometric and genetic data. Zoologischer Anzeiger 296, 129-140. DOI: 10.1016/j.jcz.2021.12.004
Oladipupo, A. A., Kelly, A. M., Davis, D. A., Bruce, T. J., 2023. Investigation of dietary exogenous protease and humic substance on growth, disease resistance to Flavobacterium covae and immune responses in juvenile channel catfish (Ictalurus punctatus). Journal of Fish Diseases pp. 1-19. DOI: 10.1111/jfd.13835
Prokešová, M., Bušová, M., Zare, M., Tran, H. Q., Kučerová, E., Ivanova, A. P., Gebauer, T., Stejskal, V., 2021. Effect of humic substances as feed additive on the growth performance, antioxidant status, and health condition of african catfish (Clarias gariepinus, Burchell 1822). Animals 11(8), 2266. DOI: 10.3390/ani11082266
Rasidi, R., Jusadi, D., Setiawati, M., Yuhana, M., Zairin Jr, M., Sugama, K., 2021. Dietary Supplementation of humic acid in the Feed of juvenile asian seabass, Lates calcarifer to counteract possible negative effects of Cadmium Accumulation on Growth and Fish Well‐being when Green Mussel (Perna viridis) is used as a Feed ingredient. Aquaculture Research 52(6), 2550-2568. DOI: 10.1111/are.15104
Steinberg, C., Timofeyev, M. A., Menzel, R., 2009. Dissolved humic substances—interactions with organisms, Academic Press Oxford: Oxford, UK.
Steinberg, C. E., Meinelt, T., Timofeyev, M. A., Bittner, M., Menzel, R., 2008. Humic substances: Part 2: Interactions with organisms. Environmental Science and Pollution Research 15(2), 128-135. DOI: 10.1065/espr2007.07.434
Thongprajukaew, K., Kovitvadhi, U., Kovitvadhi, S., Somsueb, P., Rungruangsak-Torrissen, K., 2011. Effects of different modified diets on growth, digestive enzyme activities and muscle compositions in juvenile Siamese fighting fish (Betta splendens Regan, 1910). Aquaculture 322(323), 1-9. DOI: 10.1016/j.aquaculture.2011.10.006
Torrecillas, S., Terova, G., Makol, A., Serradell, A., Valdenegro, V., Gini, E., Izquierdo, M., Acosta, F., Montero, D., 2019. Dietary phytogenics and galactomannan oligosaccharides in low fish meal and fish oil-based diets for European sea bass (Dicentrarchus labrax) juveniles: Effects on gut health and implications on in vivo gut bacterial translocation. PLoS One 14(9), e0222063. DOI: 10.1371/journal.pone.0222063
Vu, T.-D., Iwasaki, Y., Shigenobu, S., Maruko, A., Oshima, K., Iioka, E., Huang, C.-L., Abe, T., Tamaki, S., Lin, Y.-W., 2020. Behavioral and brain-transcriptomic synchronization between the two opponents of a fighting pair of the fish Betta splendens. PLoS Genetics 16(6), e1008831. DOI: 10.1371/journal.pgen.1008831
Vucskits, A., Hullár, I., Bersényi, A., Andrásofszky, E., Kulcsár, M., Szabó, J., 2010. Effect of fulvic and humic acids on performance, immune response and thyroid function in rats. Journal of Animal Physiology and Animal Nutrition 94(6), 721-728. DOI: 10.1111/j.1439-0396.2010.01023.x
Watson, C. A., DiMaggio, M., Hill, J. E., Tuckett, Q. M., Yanong, R. P., 2019. Evolution, culture, and Care for Betta splendens: FA212, 3/2019. Edis 2019(2), 1-5. DOI: 10.32473/edis-fa212-2019
Weber, J., Chen, Y., Jamroz, E., Miano, T., 2018. Preface: humic substances in the environment. Journal of Soils and Sediments 18(8), 2665-2667. DOI: 10.1007/s11368-018-2052-x
Yasar, S., Gokcimen, A., Altuntas, I., Yonden, Z., Petekkaya, E., 2002. Performance and ileal histomorphology of rats treated with humic acid preparations. Journal of Animal Physiology and Animal Nutrition 86(7‐8), 257-264. DOI: 10.1046/j.1439-0396.2002.00383.x
Yegani, M., Korver, D., 2008. Factors affecting intestinal health in poultry. Poultry Science 87(10), 2052-2063. DOI: 10.3382/ps.2008-00091
Yılmaz, S., Ergun, S., Çelik, E. Ş., Yigit, M., 2018. Effects of dietary humic acid on growth performance, haemato‐immunological and physiological responses and resistance of Rainbow trout, Oncorhynchus mykiss to Yersinia ruckeri. Aquaculture Research 49(10), 3338-3349. DOI: 10.1111/are.13798
Zhang, W., Wang, H., Brandt, D. Y., Hu, B., Sheng, J., Wang, M., Luo, H., Li, Y., Guo, S., Sheng, B., 2022. The genetic architecture of phenotypic diversity in the betta fish (Betta splendens). Science Advances 8(38), eabm4955. DOI: 10.1126/sciadv.abm495
Zoheiri, F., Hoseinifar, S. H., Mozanzadeh, M. T., Ahangarzadeh, M., Lieke, T., Van Doan, H., 2023. Dietary fulvic acid increased growth, stress tolerance and disease resistance against Vibrio harveyi in Asian seabass (Lates calcarifer) juvenile. Aquaculture Reports 32, 101738. DOI: 10.1016/j.aqrep.2023.101738
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